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Comments
Gain at zero just means don't amplify or attenuate whatever is coming in. The signal from your guitar has a level based on the peak to peak voltage output of your pickups. The interface is passing it through to the digital domain without affecting those original levels in anyway. Due to the differences in converters what that actual "number" is in digital will be different but the point of all this discussion is that you use the controls on the digital side to arrive at your in the box signal level. Or something like that!!
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But whether you use gain on the interface depends on the strength of signal from the guitar. That first stage is analog gain and it's there to get a good signal to noise ratio before the signal hits the AD convertor. Too small a signal isn't ideal, neither is near clipping.
I think most people that really know about this stuff would suggest finding a balance rather than blindly following any sort of rule. One size rarely fits all.
Btw some hard picking players with high output pickups will actually clip inputs even at zero gain so some people either recommend a pad or a DI box to get provide a pad.